WESTFIELD — The Colts defense has often made life difficult for the offense through the first week and a half of training camp.

But the mishaps suffered by the Indianapolis offense in a sloppy session on Sunday in front of a packed crowd at Grand Park were mostly self-inflicted.

A snap was botched between starting quarterback Matt Ryan and center Ryan Kelly. Stephon Gilmore picked off Ryan during 7-on-7. Nyheim Hines fumbled on a run through the middle, leading to an easy recovery and return for Gilmore.

Making matters worse, half the offense seemed to be fighting a case of the drops in the summer heat. Jonathan Taylor, Parris Campbell, Kylen Granson, Jelani Woods, Ashton Dulin and D.J. Montgomery all dropped passes in 11-on-11, and depending on the ruling, another drop could have been credited to rookie wide receiver Alec Pierce in the flat.

“That was more (drops) than we’ve had the entire training camp,” Colts head coach Frank Reich said. “What we talked about was, when you start the season, you want the arrow to just keep pointing up, up, up. It seldom does that. Today, at least offensively … too many drops, a few miscues here and there.”

The Indianapolis offense hadn’t gone through a day like Sunday’s in the first week and a half of training camp.

A star-studded Colts defense has had several big days, particularly when the team has been in full pads, but even on those days, the offense found ways to be efficient.

Ryan, who completed 11 of 18 throws Sunday, is 56 of 77 (72.7%) in 11-on-11 work in seven training camp practices, and he still hasn’t thrown an interception in full team work. The two times he’s been picked, including Gilmore’s interception Sunday, were in 7-on-7.

Indianapolis quarterbacks have thrown just three interceptions in training camp overall; none have happened in 11-on-11 practices. Wide receiver Michael Pittman Jr. has been rock solid, Dulin has been a consistent source of highlights and the vaunted Colts running game has its fair share of big plays.

But Sunday was rough overall.

“That’s part of it, that’s why we practice. We’re out here pushing ourselves to get better, to improve,” Ryan said. “We’ve had some really good days. We’ve had some days where we haven’t been quite up to the standard, but that’s how the season’s going to go. It’s not just going to be smooth sailing the whole time. Days like today are good to learn from.”

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When the Colts signed Phillip Lindsay this offseason, the veteran running back was coming off of his least productive season in the NFL, a frustrating campaign averaging 2.8 yards per carry for Miami and Houston, the two worst rushing attacks in the league.

Lindsay, a two-time 1,000-yard rusher in Denver, has looked like he’s still got plenty of ability on the practice fields at Grand Park.

Playing behind Taylor and Hines, Lindsay got some momentum going for the offense during a tough period on Sunday, ripping off a long run through the middle, catching a checkdown from Nick Foles on the next play and then turning in another nice run.

Lindsay’s initial step and quickness through the hole has been noticeable, a key development in a crowded competition for roster spots behind the team’s 1-2 punch.

With All-Pro linebacker Shaquille Leonard on the mend after offseason back surgery, veteran Zaire Franklin has been the linebacker in the starting lineup next to Bobby Okereke for most of the first-team snaps, rather than rangy backup E.J. Speed.

Franklin has spent most of his time at middle linebacker, shifting Okereke to Leonard’s spot on the weakside, and the two linebackers have made a lot of plays. Okereke broke up another pass Sunday, but it was Franklin who was most active.

Franklin broke up a pass to Taylor in 1-on-1 coverage — a drill geared heavily toward the running back — met Taylor at the goal line on another play, stopped Hines on a stretch play and pressured Ryan on another play, continuing to make his case to be in the lineup even when Leonard is back.

A couple of young defensive linemen are stacking together several impressive days in 1-on-1 pass rush drills, putting their potential on display, even if they haven’t been dominant in 11-on-11 work yet.

For example, second-year defensive end Kwity Paye has routinely given highly-respected right tackle Braden Smith trouble in 1-on-1 matchups, beating him with speed, his long arm and inside moves. Paye won both of their battles on Sunday, an indicator of how far he’s come.

Another young player who has consistently impressed in 1-on-1 drills is rookie defensive tackle Eric Johnson, a fifth-round pick with an explosive first step and remarkable quickness for a player listed a tick under 300 pounds.

Undrafted free agent JoJo Domann has been all over the field in the first week and a half of training camp.

A linebacker who plays like he has a magnet for the ball, Domann has made a handful of plays, against the pass and in the running game, where he shined while working with the second and third teams on Sunday.

Domann made at least four plays in the running game, piling up stops and continuing to make his case for a roster spot in a crowded linebacker group.

Curtis Brooks got up slowly after a big hit and left practice with trainers, but the rookie defensive tackle was only dealing with a poke in the eye, and the Colts expect him to be fine. Veteran offensive lineman Jason Spriggs, who has been playing guard in Indianapolis, also briefly left practice before returning.

Veteran safety Will Redmond was held out of practice with an undisclosed injury, joining a list of injured Colts that includes veteran tackle Dennis Kelly (knee), wide receiver DeMichael Harris, running back D’Vonte Price, safety Armani Watts, linebackers Brandon King and Forrest Rhyne and defensive tackles Chris Williams (lower right leg) and Byron Cowart.

Leonard (back) and wide receiver Michael Strachan (knee) remain on the active/physically unable to perform list.

Granson did make up for his drop a couple of plays later, coming free on a corner route for a gain of 20-plus yards. … Dayo Odeyingbo teamed with Caeveon Patton for a “sack” of Foles, added another pressure and turned in a pair of nice plays in 1-on-1 pass rush drills. … Isaiah Rodgers made a nice interception in 1-on-1 drills, running underneath Hines on a deep route and making a leaping grab. … Second-year cornerback Chris Wilcox, who was claimed off of waivers from the Buccaneers after final roster cuts last season and spent most of the year on the practice squad, also made a nice interception in 1-on-1’s, closing hard on a ball over his head. Wilcox has made a handful of nice plays in training camp so far.